The invention provides method and compositions for the modulation of
anthocyanin and proanthocyanidin production in plants. The methods of the
invention allow creation of plants having novel phenotypes. Increased
expression of anthocyanins and proanthocyanidins in plants may be used to
increase the nutritional value of food plants for both human and animal
consumption. Increased proanthocyanidin content also reduces the
potential for bloat in animals fed certain forage plants low in condensed
tannin content. The invention may also be used to modify plant
pigmentation, and for nutraceutical and food colorant production.