The present invention provides methods and compositions that allow for the
modulation of the level of at least two polynucleotides or the
polypeptides encoded thereby in a plant or plant part. Specifically, the
present invention relates to recombinant polynucleotides that are
designed to allow for the overexpression of a polynucleotide of interest
and also elicits a specific silencing effect on a second polynucleotide
of interest. In specific embodiments, the level of at least two
agronomically important sequences is modulated. Recombinant
polynucleotides capable of eliciting these effects, as well as, plants,
plant parts, seeds and grain having the recombinant polynucleotides are
also provided.