Couplers are disclosed that include first and second mutually coupled conductors. The coupled conductors may be regular or irregular in configuration, and for example, may be linear, including rectilinear or with one or more curves, bends or turns, such as forming a ring, coil, spiral or other loop. One or more sections of a coupler may be on different levels and separated by a dielectric medium, such as air or a dielectric substrate. Coupled conductors may be facing each other on the same or spaced-apart dielectric surfaces, such as opposing surfaces of a common substrate, and each conductor may include one or more portions on each side or surface of a substrate. In some examples, a coupler may include plural coupled sections, with conductors in one section being only broadside coupled, and conductors in another section being edge-coupled.