Systems of reversible USB type-A connectors which can be functionally mated with standard USB type-A connectors in each of two coupling orientations can also be reversibly mated with each other. Each plug and each receptacle includes at least one displaceable element which assumes: (a) a neutral configuration prior to coupling with the standard USB receptacle, (b) a first displaced configuration when coupled with the standard USB receptacle in a first coupling orientation, and (c) a second displaced configuration when coupled with the standard USB receptacle in a second coupling orientation. The reversible plug and the reversible receptacle are adapted for functionally mating with each other in each of two relative coupling orientations such that the displaceable element of a pre-selected one of the connectors assumes the corresponding first displaced configuration independent of the relative coupling orientation between the connectors.