A device for packaging and applying a product, which includes a container suitable for containing the product, and an application component having at least one applicator portion suitable for coming in contact with the product in order to take up some of the product. A rotational coupling arrangement is provided on at least one side of the applicator portion to allow the application component to be moved in rotation about an axis Y relative to the container. The application component includes at least one portion separating the applicator portion from the coupling means, with portion(s) having a non-constant cross section measured transversely to the axis of rotation Y.