Compositions comprising a modulated tocol content in a plant or plant part
are provided. In specific embodiments, the compositions and methods of
the invention modulate tocol content by modulating the level of a
polypeptide having a LEC1-type B domain in combination with modulating
the level of at least one other polypeptide involved in tocol
biosynthesis. Plants, plant parts, grain, seed and oil having the
modulated tocol level are also provided. Methods to enhance oxidative
stress tolerance of a plant or plant part, increase shelf-life, enhance
the nutritional value, and improve tissue quality are also provided.