A method of flowing reactants over an ion exchange membrane in a fuel cell flow field plate is provided. The flow field plate is provided, comprising a network of flow channels in the plate bounded by an electrochemically active electrode, the network comprising a series of passages having parallel grooves, the passages being interconnected by a header providing a substantially even redistribution of fluid flow received from grooves of one passage to grooves of the next passage. A reactant fluid is supplied to create a flow across the network to achieve a desired reactant utilization, wherein a flow rate and a concentration of reactant molecules per active area of membrane in the grooves increase by less than 80% across the header.