An article of footwear may have a sole structure and an upper that includes a foundation element, a strand, and a cover layer. The strand is positioned adjacent to an exterior surface of the foundation element and substantially parallel to the exterior surface for a distance of at least five centimeters. The cover layer extends along the strand for the distance of at least five centimeters, and the strand is positioned between the cover layer and the foundation element. In some configurations, the cover layer has a pair of edges on opposite sides of the strand, the strand is substantially centered between the edges of the cover layer for the distance of at least five centimeters, and areas of the exterior surface are exposed beyond the edges of the cover layer.